Karl Friston FMedSci FRSB FRS

Wellcome Principal Research Fellow and Scientific Director

Wellcome Trust Centre for Neuroimaging
Professor: Institute of Neurology, University College London
Honorary Consultant: The National Hospital for Neurology and Neurosurgery, UK




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Karl Friston is a theoretical neuroscientist and authority on brain imaging. He invented statistical parametric mapping (SPM), voxel-based morphometry (VBM) and dynamic causal modelling (DCM). These contributions were motivated by schizophrenia research and theoretical studies of value-learning, formulated as the dysconnection hypothesis of schizophrenia. Mathematical contributions include variational Laplacian procedures and generalized filtering for hierarchical Bayesian model inversion. Friston currently works on models of functional integration in the human brain and the principles that underlie neuronal interactions. His main contribution to theoretical neurobiology is a free-energy principle for action and perception (active inference). Friston received the first Young Investigators Award in Human Brain Mapping (1996) and was elected a Fellow of the Academy of Medical Sciences (1999). In 2000 he was President of the international Organization of Human Brain Mapping. In 2003 he was awarded the Minerva Golden Brain Award and was elected a Fellow of the Royal Society in 2006. In 2008 he received a Medal, College de France and an Honorary Doctorate from the University of York in 2011. He became of Fellow of the Royal Society of Biology in 2012, received the Weldon Memorial prize and Medal in 2013 for contributions to mathematical biology and was elected as a member of EMBO (excellence in the life sciences) in 2014 and the Academia Europaea in (2015). He was the 2016 recipient of the Charles Branch Award for unparalleled breakthroughs in Brain Research and the Glass Brain Award - a lifetime achievement award in the field of human brain mapping. He holds Honorary Doctorates from the University of Zurich and Radboud University.




Recent presentations


Selected Publications:





Free-energy principle and active inference



Key papers



Friston K, FitzGerald T, Rigoli F, Schwartenbeck P, Pezzulo G.

Active Inference: A Process Theory.

Neural Comput. 2016 Nov 21:1-49.


Friston K, FitzGerald T, Rigoli F, Schwartenbeck P, O'Doherty J, Pezzulo G.

Active inference and learning.

Neurosci Biobehav Rev. 2016 Sep;68:862-79.


Mirza MB, Adams RA, Mathys CD, Friston KJ.

Scene Construction, Visual Foraging, and Active Inference.

Front Comput Neurosci. 2016 Jun 14;10:56.


FitzGerald TH, Dolan RJ, Friston KJ.

Model averaging, optimal inference, and habit formation.

Front Hum Neurosci. 2014 Jun 26;8:457.


Friston K, Herreros I.

Active Inference and Learning in the Cerebellum.

Neural Comput. 2016 Sep;28(9):1812-39.


Friston K, Levin M, Sengupta B, Pezzulo G.

Knowing one's place: a free-energy approach to pattern regulation.

J R Soc Interface. 2015 Apr 6;12(105). pii: 20141383.


Friston K, Rigoli F, Ognibene D, Mathys C, Fitzgerald T, Pezzulo G.

Active inference and epistemic value.

Cogn Neurosci. 2015;6(4):187-214.


Friston KJ, Frith CD.

Active inference, communication and hermeneutics.

Cortex. 2015 Jul;68:129-43.


Friston K, Frith C.

A Duet for one.

Conscious Cogn. 2015 Nov;36:390-405.


Friston K, Schwartenbeck P, FitzGerald T, Moutoussis M, Behrens T, Dolan RJ.

The anatomy of choice: dopamine and decision-making.

Philos Trans R Soc Lond B Biol Sci. 2014 Nov 5;369(1655). pii: 20130481


Friston K, Schwartenbeck P, Fitzgerald T, Moutoussis M, Behrens T, Dolan RJ.

The anatomy of choice: active inference and agency.

Front Hum Neurosci. 2013 Sep 25;7:598.


Friston K.

Life as we know it.

J R Soc Interface. 2013 Jul 3;10(86):20130475.


Adams RA, Stephan KE, Brown HR, Frith CD, Friston KJ.

The computational anatomy of psychosis

Front Psychiatry. 2013 May 30;4:47


Brown H, Adams RA, Parees I, Edwards M, Friston K.

Active inference, sensory attenuation and illusions

Cogn Process. 2013 Jun 7. [Epub ahead of print]


Bastos AM, Usrey WM, Adams RA, Mangun GR, Fries P, Friston KJ.

Canonical microcircuits for predictive coding

Neuron. 2012 Nov 21;76(4):695-711


Friston K.

A Free Energy Principle for Biological Systems

Entropy 2012, 14, 2100-2121; doi:10.3390/e14112100


Adams RA, Shipp S, Friston KJ.

Predictions not commands: active inference in the motor system

Brain Struct Funct. 2013 May;218(3):611-43


Friston K, Samothrakis S, Montague R.

Active inference and agency: optimal control without cost functions

Biol Cybern. 2012 106(8-9):523-41


Hobson JA, Friston KJ.

Waking and dreaming consciousness: Neurobiological and functional considerations

Prog Neurobiol. 2012 May 15;98(1):82-98


Friston K, Adams RA, Perrinet L and Breakspear M

Perceptions as hypotheses: saccades as experiments

Front. Psychology (2012) 3:151. doi: 10.3389/fpsyg.2012.00151


Brown H, Friston KJ.

Free-energy and illusions: the Cornsweet effect

Front. Psychology 2012, 3:43. doi:10.3389/fpsyg.2012.00043


Friston KJ, Shiner T, FitzGerald T, Galea JM, Adams R, et al.

Dopamine, Affordance and Active Inference

PLoS Comput Biol 2012, 8(1): e1002327


Friston, K. Ao, P.

Free-energy, value and attractors

Computational and mathematical methods in medicine, Volume 2012, Article ID 937860


Friston K.

Policies and Priors

Springer Series in Computational Neuroscience, 1, Volume 10,

Computational Neuroscience of Drug Addiction, 2011;Part 3, Pages 237-283


Friston K, Mattout J, Kilner J.

Action understanding and active inference

Biol Cybern. 2011 104:137-160


Feldman H, Friston KJ.

Attention, uncertainty, and free-energy

Front Hum Neurosci. 2010 Dec 2;4:215


Friston KJ, Daunizeau J, Kilner J, Kiebel SJ.

Action and behavior: a free-energy formulation

Biol Cybern. 2010 102(3):227-60


Friston KJ, Daunizeau J, Kiebel SJ.

Reinforcement learning or active inference?

PLoS One. 2009 Jul 29;4(7):e6421


Friston K.

Hierarchical Models in the Brain

PLoS Comput Biol (2008) 4(11): e1000211


Friston K, Stephan KE.

Free energy and the brain

Synthese. 2007. 159:417-458


Friston K, Kilner J, Harrison L.

A free energy principle for the brain

J Physiol Paris. 2006 Jul-Sep;100(1-3):70-87


Friston K.
A theory of cortical responses
Philos Trans R Soc Lond B Biol Sci. 2005 Apr 29;360(1456):815-36




Reviews and perspectives


Friston K, Buzsaki G.

The Functional Anatomy of Time: What and When in the Brain.

Trends Cogn Sci. 2016 Jul;20(7):500-11.


Hobson JA, Friston K,

Consciousness, Dreams, and Inference: The Cartesian Theatre Revisited.

Journal of Consciousness Studies, 2014 21(1-2): 6-32


Hobson JA, Hong CC, Friston KJ.

Virtual reality and consciousness inference in dreaming.

Front Psychol. 2014 Oct 9;5:1133.


Quattrocki E, Friston K.

Autism, oxytocin and interoception.

Neurosci Biobehav Rev. 2014 Nov;47:410-30.


Seth AK.

Interoceptive inference, emotion, and the embodied self.

Trends Cogn Sci. 2013 Nov;17(11):565-73.


Shipp S, Adams RA, Friston KJ.

Reflections on agranular architecture: predictive coding in the motor cortex.

Trends Neurosci. 2013 Oct 21. doi:pii: S0166-2236(13)00160-4.


Gu X, Hof PR, Friston KJ, Fan J.

Anterior insular cortex and emotional awareness.

J Comp Neurol. 2013 Jun 8. doi: 10.1002/cne.23368


Sengupta B, Stemmler MB, Friston KJ.

Information and efficiency in the nervous system-a synthesis.

PLoS Comput Biol. 2013 Jul;9(7):e1003157.


Friston K,

The fantastic organ

Brain 2013: 136; 1328-1332


Friston K,

I am therefore I think

In M. Leuzinger-Bohleber, S. Arnold & M. Solms (Eds.), The unconscious: a bridge between psychoanalysis and cognitive neuroscience (pp. 113-137). London: Routledge.


Clark A.

Whatever next? Predictive brains, situated agents, and the future of cognitive science

Behav Brain Sci. 2013 Jun;36(3):181-204


Friston K, Adams R and Montague R.

What is value-accumulated reward or evidence?

Front. Neurorobot. (2012) 6:11. doi: 10.3389/fnbot.2012.00011


Edwards MJ, Adams RA, Brown H, Parees I, Friston KJ.

A Bayesian account of 'hysteria'

Brain. 2012 135(Pt 11):3495-512


Friston K.

What is optimal about motor control?

Neuron. 2011 Nov 3;72(3):488-98


Friston K.

The free-energy principle: a unified brain theory?

Nat Rev Neurosci. 2010 Feb;11(2):127-38


Friston K.

The free-energy principle: a rough guide to the brain?

Trends Cogn Sci. 2009 13(7):293-301


Friston K.

Embodied Inference: or "I think therefore I am, if I am what I think"

In: The Implications of Embodiment (Cognition and Communication) Eds. Wolfgang Tschacher and Claudia Bergomi

Imprint Academic ISBN 9781845402402



Technical papers


Sengupta B, Tozzi A, Cooray GK, Douglas PK, Friston KJ.

Towards a Neuronal Gauge Theory.

PLoS Biol. 2016 Mar 8;14(3):e1002400.


Friston K, Breakspear M and Deco G

Perception and self-organized instability

Front. Comput. Neurosci. (2012) 6:44. doi: 10.3389/fncom.2012.00044


Friston K, Stephan K, Li B, and Daunizeau J.

Generalised Filtering

Mathematical Problems in Engineering. vol. 2010, Article ID 621670, 34 pages, 2010


Friston KJ.

Variational filtering

Neuroimage. 2008 Jul 1;41(3):747-66


Friston KJ, Trujillo-Barreto N, Daunizeau J.

DEM: A variational treatment of dynamic systems

Neuroimage. 2008 Jul 1;41(3):849-85


Friston K, Mattout J, Trujillo-Barreto N, Ashburner J, Penny W.

Variational free energy and the Laplace approximation

NeuroImage. 2007 Jan 1;34(1):220-34



Features, commentaries and popular science


Hobson, JA, Friston, KJ

A Response to Our Theatre Critics

JOURNAL OF CONSCIOUSNESS STUDIES. 2016  23 (3-4) pp. 245-254.


Friston KJ, Lawson R, Frith CD.

On hyperpriors and hypopriors: comment on Pellicano and Burr

Trends Cogn Sci. 2013 Jan;17(1):1.


Friston K.

Competitive dynamics in the brain: Comment on "Information flow dynamics in the brain" by M.I. Rabinovich et al.

Phys Life Rev. 2012 Mar;9(1):76-7.


Friston K,

Predictive coding, precision and synchrony

Cognitive Neuroscience. 2012. Volume 3, Issue 3-4:238-239.


Friston K, Thornton C, Clark A.

Free-energy minimization and the dark-room problem.

Front Psychol. 2012;3:130.- pdf


Friston K.

Prediction, perception and agency.

Int J Psychophysiol. 2012. 83:248-252 - pdf


Friston K.

The history of the future of the Bayesian brain.

Neuroimage. 2011 Oct 17- pdf


Is this a unified theory of the brain? From New Scientist. 28 May 2008 pp 30-33- pdf

By Gregory T. Huang


The Prophetic Brain: Universe in 2009 SEED Magazine- pdf

By Karl Friston / January 27, 2009


Surprise yourself. The Social Brain RSA. July 15, 2009 - pdf

By Matt Grist


Thornton C.

Some puzzles relating to the free-energy principle: comment on Friston.

Trends Cogn Sci. 2010 Feb;14(2):53-4; author reply 54-5.- pdf


Friston K.

Is the free-energy principle neurocentric?

Nat Rev Neurosci. 2010 11:605 - pdf


McCrone, J.

Wild minds, New scientist, 2112, 3 Dec, 1997. p 26-30




http://assets0.serious-science.org/style/logo.png Free Energy Principle

Neuroscientist Karl Friston on the Markov blanket, Bayesian model evidence, and different global brain theories




TV Interview (Spanish and English)

Portada de RTVE.es


Redes - La fórmula del cerebro (V.O.)
released: 20 November 2011

Lecture (for schoolchildren)


 Image result for european parliament

Brain Awareness Week at the European Parliament
published: March. 19, 2010, recorded: March 2008

Free-energy and the brain- (PowerPoint)


Lectures (for grownups)


event thumbnail image



Free energy and active inference


Karl Friston, Wellcome Trust Centre for Neuroimaging, University College London

published: Oct. 16, 2012, recorded: September 2012





Embodied inference and free energy


Karl Friston, Wellcome Trust Centre for Neuroimaging, University College London

published: Nov 2013, recorded: March 2013



Theoretical neurobiology


Sengupta B, Stemmler MB, Friston KJ.

Information and efficiency in the nervous system--a synthesis.

PLoS Comput Biol. 2013 Jul;9(7):e1003157.


Kiebel SJ, Friston KJ.

Free energy and dendritic self-organization.

Front Syst Neurosci. 2011;5:80.


Brown H, Friston K, Bestamnn S.

Active inference, attention and motor preparation.

Front Pyshcology2:218. doi: 10.3389/fpsyg.2011.00218- pdf


Kiebel SJ, von Kriegstein K, Daunizeau J, Friston KJ.

Recognizing sequences of sequences.

PLoS Comput Biol. 2009 Aug;5(8):e1000464. - pdf


Friston K.

Learning and inference in the brain.

Neural Netw. 2003 Nov;16(9):1325-52.- pdf


Friston K.
Functional integration and inference in the brain.
Prog Neurobiol. 2002 Oct;68(2):113-43.- pdf


Friston KJ.
Brain function nonlinear coupling and neuronal transients. 
Neuroscientist. 2001 Oct;7(5):406-18.- pdf
Friston KJ, Price CJ. 
Dynamic representations and generative models of brain function.
Brain Res Bull. 2001 Feb;54(3):275-85. - pdf
Friston KJ.
The labile brain. I. Neuronal transients and nonlinear coupling. 
Philos Trans R Soc Lond B Biol Sci. 2000 Feb 29;355(1394):215-36.- pdf
Friston KJ.
The labile brain. II. Transients, complexity and selection. 
Philos Trans R Soc Lond B Biol Sci. 2000 Feb 29;355(1394):237-52.- pdf
Friston KJ.
The labile brain. III. Transients and spatio-temporal receptive fields. 
Philos Trans R Soc Lond B Biol Sci. 2000 Feb 29;355(1394):253-65.- pdf
Friston KJ.
Another neural code? 
NeuroImage. 1997 Apr;5(3):213-20.- pdf
Friston KJ.
Transients, metastability, and neuronal dynamics. 
NeuroImage. 1997 Feb;5(2):164-71.- pdf
Friston KJ.
Neuronal transients. 
Proc Biol Sci. 1995 Sep 22;261(1362):401-5.- pdf
Friston, K. J., Tononi, G., Sporns, O. and Edelman, G. M. 
Characterising the complexity of neuronal interactions. 
Hum. Brain Mapp., 1995 3: 302-314.- pdf



Predictive coding and information theory


Auksztulewicz R, Friston K.

Attentional Enhancement of Auditory Mismatch Responses: a DCM/MEG Study.

Cereb Cortex. 2015 Nov;25(11):4273-83.


Auksztulewicz R, Friston K.

Repetition suppression and its contextual determinants in predictive coding.

Cortex. 2016 Jul;80:125-40.


Kanai R, Komura Y, Shipp S, Friston K.

Cerebral hierarchies: predictive processing, precision and the pulvinar.

Philos Trans R Soc Lond B Biol Sci. 2015 May 19;370(1668).


Bastos AM, Litvak V, Moran R, Bosman CA, Fries P, Friston KJ.

A DCM study of spectral asymmetries in feedforward and feedback connections between visual areas V1 and V4 in the monkey.

Neuroimage. 2015 Mar;108:460-75.


Brown HR, Friston KJ.

The functional anatomy of attention: a DCM study.

Front Hum Neurosci. 2013 Dec 2;7:784.


Perrinet LU, Adams RA, Friston KJ.

Active inference, eye movements and oculomotor delays.

Biol Cybern. 2014 Dec;108(6):777-801.


Brown HR, Friston KJ.

Dynamic causal modelling of precision and synaptic gain in visual perception - an EEG study.

Neuroimage. 2012 Oct 15;63(1):223-31.


Coste CP, Sadaghiani S, Friston KJ, and Kleinschmidt A.

Ongoing Brain Activity Fluctuations Directly Account for Intertrial and Indirectly for Intersubject Variability in Stroop Task Performance

Cerebral Cortex doi:10.1093/cercor/bhr050 - pdf


Hesselmann G, Sadaghiani S, Friston KJ, Kleinschmidt A.

Predictive coding or evidence accumulation? False inference and neuronal fluctuations.

PLoS One. 2010 Mar 29;5(3):e9926 - pdf


Friston KJ, Kiebel SJ.

Attractors in song.

New Math. Nat. Comput. 2009 5:83-114.- pdf


Friston KJ, Kiebel SJ.

Cortical circuits for perceptual inference.

Neural Netw. 2009 Jul 19. [Epub ahead of print]- pdf


Friston KJ, Kiebel SJ.

Predictive coding under the free-energy principle

Phil. Trans. R. Soc. B May 12, 2009 364:1211-1221- pdf


Brodersen KH, Penny WD, Harrison LM, Daunizeau J, Ruff CC, Duzel E, Friston KJ, Stephan KE.

Integrated Bayesian models of learning and decision making for saccadic eye movements.

Neural Netw. 2008 Nov;21(9):1247-60.- pdf


Garrido MI, Kilner JM, Kiebel SJ, Friston KJ.

Evoked brain responses are generated by feedback loops.

Proc Natl Acad Sci U S A. 2007 Dec 26;104(52):20961-6.- pdf


Furl N, van Rijsbergen NJ, Treves A, Friston KJ, Dolan RJ.

Experience-dependent coding of facial expression in superior temporal sulcus.

Proc Natl Acad Sci U S A. 2007 Aug 7;- pdf


Griffiths TD, Kumar S, Warren JD, Stewart L, Stephan KE, Friston KJ.

Approaches to the cortical analysis of auditory objects.

Hear Res. 2007 Jul;229(1-2):46-53. Epub 2007 Jan 16.- pdf


Kilner JM, Friston KJ, Frith CD.

Predictive coding: an account of the mirror neuron system.

Cogn Process. 2007 Sep;8(3):159-66.- pdf


Kilner JM, Friston KJ, Frith CD.

The mirror-neuron system: a Bayesian perspective.

Neuroreport. 2007 Apr 16;18(6):619-23.


Harrison LM, Stephan KE, Rees G, Friston KJ.

Extra-classical receptive field effects measured in striate cortex with fMRI.

NeuroImage. 2007 Feb 1;34(3):1199-208.- pdf


Harrison LM, Duggins A, Friston KJ.

Encoding uncertainty in the hippocampus.

Neural Netw. 2006 Jun;19(5):535-46.- pdf


Strange BA, Duggins A, Penny W, Dolan RJ, Friston KJ.
Information theory, novelty and hippocampal responses: unpredicted or unpredictable? 
Neural Netw. 2005 Apr;18(3):225-30.- pdf



Computational neuroscience



Schwartenbeck P, FitzGerald TH, Mathys C, Dolan R, Kronbichler M, Friston K.

Evidence for surprise minimization over value maximization in choice behavior.

Sci Rep. 2015 Nov 13;5:16575.


Marreiros AC, Kiebel SJ, Daunizeau J, Harrison LM, Friston KJ.

Population dynamics under the Laplace assumption.

Neuroimage. 2009 Feb 1;44(3):701-14.


Marreiros AC, Daunizeau J, Kiebel SJ, Friston KJ.

Population dynamics: Variance and the sigmoid activation function.

NeuroImage. 2008 Apr 29.


Chumbley JR, Dolan RJ, Friston KJ.

Attractor models of working memory and their modulation by reward.

Biol Cybern. 2008 Jan;98(1):11-8.- pdf


David O, Kilner JM, Friston KJ.

Mechanisms of evoked and induced responses in MEG/EEG.

NeuroImage. 2006 Jul 15;31(4):1580-91.- pdf


Harrison LM, David O, Friston KJ. 
Stochastic models of neuronal dynamics. 
Philos Trans R Soc Lond B Biol Sci. 2005 May 29;360(1457):1075-91.- pdf


David O, Harrison L, Friston KJ. 
Modelling event-related responses in the brain. 
NeuroImage. 2005 Apr 15;25(3):756-70.- pdf
David O, Cosmelli D, Friston KJ.
Evaluation of different measures of functional connectivity using a neural mass model. 
NeuroImage. 2004 Feb;21(2):659-73.- pdf
Breakspear M, Terry JR, Friston KJ.
Modulation of excitatory synaptic coupling facilitates synchronization and complex dynamics in a biophysical model of neuronal dynamics. 
Network. 2003 Nov;14(4):703-32.- pdf
David O, Friston KJ. 
A neural mass model for MEG/EEG: coupling and neuronal dynamics. 
NeuroImage. 2003 Nov;20(3):1743-55.- pdf
Chawla D, Friston KJ, Lumer ED.
Zero-lag synchronous dynamics in triplets of interconnected cortical areas. 
Neural Netw. 2001 Jul-Sep;14(6-7):727-35.- pdf

Chawla D, Lumer ED, Friston KJ

Relating macroscopic measures of brain activity to fast, dynamic neuronal interactions

Neural Computation 12, 2805-2821 (200) - pdf

Rees G, Friston K, Koch C.
A direct quantitative relationship between the functional properties of human and macaque V5. 
Nat Neurosci. 2000 Jul;3(7):716-23.- (pdf)
Chawla D, Lumer ED, Friston KJ.
The relationship between synchronization among neuronal populations and their mean activity levels. 

Neural Comput. 1999 Aug 15;11(6):1389-411. - pdf

Friston KJ Stephan KM Frackowiak RSJ
Transient phase-locking and dynamic correlations: Are they the same thing?
Human Brain Mapping1997;5:48-57. - pdf
Friston KJ, Frith CD, Frackowiak RS. 
Principal component analysis learning algorithms: a neurobiological analysis. 
Proc Biol Sci. 1993 Oct 22;254(1339):47-54. - pdf
Friston KJ, Frith CD, Liddle PF, Frackowiak RS.
Investigating a network model of word generation with positron emission tomography. 
Proc Biol Sci. 1991 May 22;244(1310):101-6. - pdf


Computational psychiatry



Schwartenbeck P, Friston K.

Computational Phenotyping in Psychiatry: A Worked Example.

eNeuro. 2016 Aug 2;3(4).


Friston KJ, Stephan KE, Montague R, Dolan RJ

Computational psychiatry: the brain as a phantastic organ

The Lancet Psychiatry, 1(2),148-158


Picard F, Friston K.

Predictions, perception, and a sense of self.

Neurology. 2014 Sep 16;83(12):1112-8.


Lawson RP, Rees G, Friston KJ.

An aberrant precision account of autism.

Front Hum Neurosci. 2014 May 14;8:302.


Dynamic Causal Modelling


Bastos AM, Litvak V, Moran R, Bosman CA, Fries P, Friston KJ.

A DCM study of spectral asymmetries in feedforward and feedback connections between visual areas V1 and V4 in the monkey.

Neuroimage. 2015 Mar;108:460-75.


Adams RA, Aponte E, Marshall L, Friston KJ.

Active inference and oculomotor pursuit: the dynamic causal modelling of eye movements.

J Neurosci Methods. 2015 Mar 15;242:1-14.


Cooray GK, Sengupta B, Douglas P, Englund M, Wickstrom R, Friston K.

Characterising seizures in anti-NMDA-receptor encephalitis with dynamic causal modelling. Neuroimage. 2015 Sep;118:508-19.


Papadopoulou M, Leite M, van Mierlo P, Vonck K, Lemieux L, Friston K, Marinazzo D.

Tracking slow modulations in synaptic gain using dynamic causal modelling: validation in epilepsy.

Neuroimage. 2015 Feb 15;107:117-26.


Razi A, Kahan J, Rees G, Friston KJ.

Construct validation of a DCM for resting state fMRI.

Neuroimage. 2015 Feb 1;106:1-14.


Friston KJ, Kahan J, Biswal B, Razi A.

A DCM for resting state fMRI.

Neuroimage. 2014 Jul 1;94:396-407.


Friston KJ, Bastos AM, Oswal A, van Wijk B, Richter C, Litvak V.

Granger causality revisited.

Neuroimage. 2014 Nov 1;101:796-808.


Friston KJ, Kahan J, Razi A, Stephan KE, Sporns O.

On nodes and modes in resting state fMRI.

Neuroimage. 2014 Oct 1;99:533-47.


Pinotsis DA, Brunet N, Bastos A, Bosman CA, Litvak V, Fries P, Friston KJ.

Contrast gain control and horizontal interactions in V1: a DCM study.

Neuroimage. 2014 May 15;92:143-55.


Seghier ML, Friston KJ.

Network discovery with large DCMs.

Neuroimage. 2012 68 (2013) 181-191. - pdf


Pinotsis DA, Moran RJ, Friston KJ.

Dynamic causal modeling with neural fields.

Neuroimage. 2012 Jan 16;59(2):1261-74. - pdf


Friston KJ, Bastos A, Litvak V, Stephan KE, Fries P, Moran RJ.

DCM for complex-valued data: cross-spectra, coherence and phase-delays.

Neuroimage. 2012 Jan 2;59(1):439-55. - pdf


Moran RJ, Stephan KE, Dolan RJ, Friston KJ.

Consistent spectral predictors for dynamic causal models of steady-state responses.

Neuroimage. 2011 Apr 15;55(4):1694-708. - pdf


Havlicek M, Friston KJ, Jan J, Brazdil M, Calhoun VD.

Dynamic modeling of neuronal responses in fMRI using cubature Kalman filtering.

Neuroimage. 2011 Jun 15;56(4):2109-28. - pdf


Friston KJ, Li B, Daunizeau J, Stephan KE.

Network discovery with DCM.

Neuroimage. 2010 Dec 21. - pdf


Li B, Daunizeau J, Stephan KE, Penny W, Hu D, Friston K.

Generalised filtering and stochastic DCM for fMRI.

Neuroimage. 2011 Feb 16. - pdf


Marreiros AC, Kiebel SJ, Friston KJ.

A dynamic causal model study of neuronal population dynamics.

Neuroimage. 2010 May 15;51(1):91-101.- pdf


Stephan KE, Penny WD, Moran RJ, den Ouden HE, Daunizeau J, Friston KJ.

Ten simple rules for dynamic causal modeling.

Neuroimage. 2010 Feb 15;49(4):3099-109. - pdf


Penny WD, Litvak V, Fuentemilla L, Duzel E, Friston K.

Dynamic Causal Models for phase coupling.

J Neurosci Methods. 2009 Sep 30;183(1):19-30. - pdf


Daunizeau J, Kiebel SJ, Friston KJ.

Dynamic causal modelling of distributed electromagnetic responses.

Neuroimage. 2009 Aug 15;47(2):590-601. - pdf


Chen CC, Henson RN, Stephan KE, Kilner JM, Friston KJ.

Forward and backward connections in the brain: a DCM study of functional asymmetries.

Neuroimage. 2009 Apr 1;45(2):453-62 - pdf


Moran RJ, Stephan KE, Seidenbecher T, Pape HC, Dolan RJ, Friston KJ.

Dynamic causal models of steady-state responses.

Neuroimage. 2009 Feb 1;44(3):796-811. - pdf


Stephan KE, Kasper L, Harrison LM, Daunizeau J, den Ouden HE, Breakspear M, Friston KJ.

Nonlinear dynamic causal models for fMRI.

Neuroimage. 2008 May 11 - pdf


Moran RJ, Stephan KE, Kiebel SJ, Rombach N, O'Connor WT, Murphy KJ, Reilly RB, Friston KJ.

Bayesian estimation of synaptic physiology from the spectral responses of neural masses.

Neuroimage. 2008 Feb 1. [Epub ahead of print] - pdf


Chen CC, Kiebel SJ, Friston KJ.

Dynamic causal modelling of induced responses.

Neuroimage. 2008 Jul 15;41(4):1293-312. - pdf


Marreiros AC, Kiebel SJ, Friston KJ.

Dynamic causal modelling for fMRI: a two-state model.

Neuroimage. 2008 Jan 1;39(1):269-78. - pdf


Stephan KE, Weiskopf N, Drysdale PM, Robinson PA, Friston KJ.

Comparing hemodynamic models with DCM.

NeuroImage. 2007 Nov 15;38(3):387-401 - pdf


Garrido MI, Kilner JM, Kiebel SJ, Stephan KE, Friston KJ.

Dynamic causal modelling of evoked potentials: A reproducibility study.

NeuroImage. 2007 Mar 27; [Epub ahead of print]- pdf


Kiebel SJ, Garrido MI, Friston KJ.

Dynamic causal modelling of evoked responses: The role of intrinsic connections.

NeuroImage. 2007 Jun;36(2):332-45.- pdf


Stephan KE, Harrison LM, Kiebel SJ, David O, Penny WD, Friston KJ.

Dynamic causal models of neural system dynamics: current state and future extensions.

J Biosci. 2007 Jan;32(1):129-44.


Kiebel SJ, Kloppel S, Weiskopf N, Friston KJ.

Dynamic causal modeling: a generative model of slice timing in fMRI.

NeuroImage. 2007 Feb 15;34(4):1487-96.- pdf


Mouridsen K, Friston K, Hjort N, Gyldensted L, Ostergaard L, Kiebel S.

Bayesian estimation of cerebral perfusion using a physiological model of microvasculature.

NeuroImage. 2006 Nov 1;33(2):570-9.- pdf


Kiebel SJ, David O, Friston KJ.

Dynamic causal modelling of evoked responses in EEG/MEG with lead field parameterization.

NeuroImage. 2006 May 1;30(4):1273-84.- pdf


David O, Kiebel SJ, Harrison LM, Mattout J, Kilner JM, Friston KJ.

Dynamic causal modeling of evoked responses in EEG and MEG.

NeuroImage. 2006 May 1;30(4):1255-72.- pdf


Stephan KE, Penny WD, Marshall JC, Fink GR, Friston KJ.

Investigating the functional role of callosal connections with dynamic causal models.

Ann N Y Acad Sci. 2005 Dec;1064:16-36.


Lee L, Friston K, Horwitz B.

Large-scale neural models and dynamic causal modelling.

NeuroImage. 2006 May 1;30(4):1243-54.- pdf


Penny WD, Stephan KE, Mechelli A, Friston KJ.
Comparing dynamic causal models.
NeuroImage. 2004 Jul;22(3):1157-72.- pdf


Penny WD, Stephan KE, Mechelli A, Friston KJ.
Modelling functional integration: a comparison of structural equation and dynamic causal models. 
NeuroImage. 2004;23 Suppl 1:S264-74.- pdf


Mechelli A, Price CJ, Noppeney U, Friston KJ.
A dynamic causal modeling study on category effects: bottom-up or top-down mediation? 
J Cogn Neurosci. 2003 Oct 1;15(7):925-34. 


Friston KJ, Harrison L, Penny W.
Dynamic causal modelling. 
NeuroImage. 2003 Aug;19(4):1273-302.- pdf

Empirical Bayes and hierarchical model inversion


Friston K, Zeidman P, Litvak V.

Empirical Bayes for DCM: A Group Inversion Scheme.

Front Syst Neurosci. 2015 Nov 27;9:164.


Friston KJ, Litvak V, Oswal A, Razi A, Stephan KE, van Wijk BC, Ziegler G, Zeidman P.

Bayesian model reduction and empirical Bayes for group (DCM) studies.

Neuroimage. 2016 Mar;128:413-31. air


Bayesian and Variational model inversion



Balaji B, and Friston K.

Bayesian state estimation using generalized coordinates.

Proc. SPIE 8050, 80501Y (2011)- pdf


Friston K, Penny W.

Post hoc Bayesian model selection.

Neuroimage. 2011 Jun 15;56(4):2089-99. - pdf


Friston K, Stephan K, Li B, and Daunizeau J.

Generalised Filtering

Mathematical Problems in Engineering. vol. 2010, Article ID 621670, 34 pages, 2010.- pdf


Friston KJ.

Variational filtering.

Neuroimage. 2008 Jul 1;41(3):747-66.- pdf


Friston KJ, Trujillo-Barreto N, Daunizeau J.

DEM: A variational treatment of dynamic systems.

Neuroimage. 2008 Jul 1;41(3):849-85.- pdf


Friston K, Mattout J, Trujillo-Barreto N, Ashburner J, Penny W.

Variational free energy and the Laplace approximation.

NeuroImage. 2007 Jan 1;34(1):220-34.- pdf


Penny W, Ghahramani Z, Friston K 
Bilinear dynamical systems. 
Philos Trans R Soc Lond B Biol Sci. 2005 May 29;360(1457):983-93.- pdf
Penny WD, Trujillo-Barreto NJ, Friston KJ.
Bayesian fMRI time series analysis with spatial priors. 
NeuroImage. 2005 Jan 15;24(2):350-62.- pdf
Harrison L, Penny WD, Friston K.
Multivariate autoregressive modeling of fMRI time series. 
NeuroImage. 2003 Aug;19(4):1477-91.- pdf
Penny W, Kiebel S, Friston K.
Variational Bayesian inference for fMRI time series. 
NeuroImage. 2003 Jul;19(3):727-41.- pdf 

Friston KJ, Penny W

Posterior probability maps and SPMs

NeuroImage, (2003) 19, 1240-1249.- pdf

Phillips C, Rugg MD, Friston KJ.
Anatomically informed basis functions for EEG source localization: combining functional and anatomical constraints. 
NeuroImage. 2002 Jul;16(3 Pt 1):678-95.- pdf

Friston KJ

Bayesian Estimation of Dynamical Systems: An Application to fMRI

NeuroImage 16, 513-530 (2002)- pdf


Friston KJ, Penny W, Phillips, Kiebel S, Hinton G, Ashburner J

Classical and Bayesian Inference in Neuroimaging: Theory

NeuroImage 16, 465-483 (2002)- pdf


Friston KJ, Glaser DE, Henson RNA, Kiebel S, Phillips C, Ashburner J

Classical and Bayesian Inference in Neuroimaging: Applications

NeuroImage 16, 484-512 (2002)- pdf

Phillips C, Rugg MD & Friston KJ.
Systematic Regularization of Linear Inverse Solutions of the EEG Source Localization Problem. 
NeuroImage 2002;17:287-301- pdf
Büchel C, Friston KJ.
Dynamic changes in effective connectivity characterized by variable parameter regression and Kalman filtering. 
Hum Brain Mapp. 1998;6(5-6):403-8.- pdf



 event thumbnail image

Variational filtering in generated coordinates of motion
published: Sept. 9, 2008, recorded: May 2008

Variational filtering and DEM- (PowerPoint)


Optimality, value-learning and plasticity



Mathys C, Jean Daunizeau J, Karl J Friston KJ, Klaas Enno Stephan KE.

A Bayesian foundation for individual learning under uncertainty

Front. Hum. Neurosci. 5:35 DOI: 10.3389/fnhum.2011.00039 - pdf


Daunizeau J, den Ouden HE, Pessiglione M, Kiebel SJ, Stephan KE, Friston KJ.

Observing the observer (I): meta-bayesian models of learning and decision-making.

PLoS One. 2010 Dec 14;5(12):e15554. - pdf


Daunizeau J, den Ouden HE, Pessiglione M, Kiebel SJ, Friston KJ, Stephan KE.

Observing the observer (II): deciding when to decide.

PLoS One. 2010 Dec 14;5(12):e15555. - pdf


Garrido MI, Kilner JM, Kiebel SJ, Stephan KE, Baldeweg T, Friston KJ.

Repetition suppression and plasticity in the human brain.

Neuroimage. 2009 Oct 15;48(1):269-79.- pdf


Yoshida W, Dolan RJ, Friston KJ.
Game theory of mind.
PLoS Comput Biol. 2008 Dec;4(12):e1000254.- pdf
Seymour B, O'Doherty JP, Dayan P, Koltzenburg M, Jones AK, Dolan RJ, Friston KJ, Frackowiak RS.
Temporal difference models describe higher-order learning in humans. 
Nature. 2004 Jun 10;429(6992):664-7.- pdf
O'Doherty J, Dayan P, Schultz J, Deichmann R, Friston K, Dolan RJ.
Dissociable roles of ventral and dorsal striatum in instrumental conditioning. 
Science. 2004 Apr 16;304(5669):452-4. 
O'Doherty JP, Dayan P, Friston K, Critchley H, Dolan RJ.
Temporal difference models and reward-related learning in the human brain. 
Neuron. 2003 Apr 24;38(2):329-37.- pdf
Thiel CM, Friston KJ, Dolan RJ.
Cholinergic modulation of experience-dependent plasticity in human auditory cortex. 
Neuron. 2002 Aug 1;35(3):567-74.- pdf
Elliott R, Friston KJ, Dolan RJ.
Dissociable neural responses in human reward systems. 
J Neurosci. 2000 Aug 15;20(16):6159-65. 
Büchel C, Coull JT, Friston KJ.
The predictive value of changes in effective connectivity for human learning. 
Science. 1999 Mar 5;283(5407):1538-41.- pdf
Friston KJ, Tononi G, Reeke GN Jr, Sporns O, Edelman GM.
Value-dependent selection in the brain: simulation in a synthetic neural model. 
Neuroscience. 1994 Mar;59(2):229-43.- pdf



Dysconnection hypothesis and schizophrenia



Friston K, Brown HR, Siemerkus J, Stephan KE.

The dysconnection hypothesis (2016).

Schizophr Res. 2016 Oct;176(2-3):83-94.


Friston K.

The Bayesian Savant.

Biol Psychiatry. 2016 Jul 15;80(2):87-9.


Ranlund S, Adams RA, Díez Á, Constante M, Dutt A, Hall MH, Maestro Carbayo A, McDonald C, Petrella S, Schulze K, Shaikh M, Walshe M, Friston K, Pinotsis D, Bramon E.

Impaired prefrontal synaptic gain in people with psychosis and their relatives during the mismatch negativity.

Hum Brain Mapp. 2016 Jan;37(1):351-65.


Bastos-Leite AJ, Ridgway GR, Silveira C, Norton A, Reis S, Friston KJ.

Dysconnectivity within the default mode in first-episode schizophrenia: a stochastic dynamic causal modeling study with functional magnetic resonance imaging.

Schizophr Bull. 2015 Jan;41(1):144-53.


Fogelson N, Litvak V, Peled A, Fernandez-del-Olmo M, Friston K.

The functional anatomy of schizophrenia: A dynamic causal modeling study of predictive coding.

Schizophr Res. 2014 Sep;158(1-3):204-12.


Adams RA, Perrinet LU, Friston K.

Smooth pursuit and visual occlusion: active inference and oculomotor control in schizophrenia.

PLoS One. 2012;7(10):e47502.


Stephan KE, Karl J. Friston KJ, Frith CD

Dysconnection in Schizophrenia: From Abnormal Synaptic Plasticity to Failures of Self-monitoring

Schizophrenia Bulletin 2009; doi: 10.1093/schbul/sbn176.


Stephan KE, Baldeweg T, Friston KJ.

Synaptic plasticity and dysconnection in schizophrenia.

Biol Psychiatry. 2006 May 15;59(10):929-39. Epub 2006 Jan 19. Review


Friston KJ.

Hallucinations and perceptual inference

Behavioral and Brain Sciences [0140-525X]: 2005 28(6):764-766- pdf


Friston KJ
Disconnection and cognitive dysmetria in schizophrenia. 
Am J Psychiatry. 2005 Mar;162(3):429-32.- pdf


Friston KJ.
Dysfunctional connectivity in schizophrenia. 
World Psychiatry. 2002 Jun;1(2):66-71.- pdf
Friston KJ.
Schizophrenia and the disconnection hypothesis. 
Acta Psychiatr Scand Suppl. 1999;395:68-79. 
Friston KJ.
The disconnection hypothesis. 
Schizophr Res. 1998 Mar 10;30(2):115-25.- pdf
Friston KJ.
Theoretical neurobiology and schizophrenia. 
Br Med Bull. 1996 Jul;52(3):644-55. - pdf


Friston KJ, Frith CD.
Schizophrenia: a disconnection syndrome? 
Clin Neurosci. 1995;3(2):89-97.- pdf







Noppeney U, Friston KJ, Price CJ.
Degenerate neuronal systems sustaining cognitive functions. 
J Anat. 2004 Dec;205(6):433-42.
Friston KJ, Price CJ.
Degeneracy and redundancy in cognitive anatomy. 
Trends Cogn Sci. 2003 Apr;7(4):151-152. - pdf
Price CJ, Friston KJ.
Degeneracy and cognitive anatomy. 
Trends Cogn Sci. 2002 Oct 1;6(10):416-421. - pdf
Price CJ, Mummery CJ, Moore CJ, Frackowiak RS, Friston KJ. 
Delineating necessary and sufficient neural systems with functional imaging studies of neuropsychological patients. 
J Cogn Neurosci. 1999 Jul;11(4):371-82. 



Neuroimaging (reviews and commentaries)


Friston K, Moran R, Seth AK.

Analysing connectivity with Granger causality and dynamic causal modelling.

Curr Opin Neurobiol. 2012 Dec 21. doi:pii: S0959-4388(12)00184-5. - pdf


Friston K.

Ten ironic rules for non-statistical reviewers.

Neuroimage. 2012 61: 1300-1310- pdf


Friston KJ.

Functional and Effective Connectivity: A Review

Brain Connectivity. 2011, 1(1): 13-36.- pdf


Friston KJ, Dolan RJ.

Computational and dynamic models in neuroimaging.

Neuroimage. 2009 Dec 28. [Epub ahead of print]- pdf


Friston KJ.

Modalities, modes, and models in functional neuroimaging.

Science. 2009 Oct 16;326(5951):399-403.- pdf


Kiebel SJ, Garrido MI, Moran R, Chen CC, Friston KJ.

Dynamic causal modeling for EEG and MEG.

Hum Brain Mapp. 2009 Jun;30(6):1866-76. Review.- pdf


Friston K.

Causal modelling and brain connectivity in functional magnetic resonance imaging.

PLoS Biol. 2009 Feb 17;7(2):e33.- pdf


Friston KJ, Rotshtein P, Geng JJ, Sterzer P, Henson RN.

A critique of functional localisers.

NeuroImage. 2006 May 1;30(4):1077-87.- pdf


Friston KJ.

Models of brain function in neuroimaging.

Annu Rev Psychol. 2005;56:57-87- pdf


Friston K. 
Beyond phrenology: what can neuroimaging tell us about distributed circuitry? 
Annu Rev Neurosci. 2002;25:221-50. Epub 2002 Mar 19.- pdf
Friston KJ.
Imaging neuroscience: principles or maps?
Proc Natl Acad Sci U S A. 1998 Feb 3;95(3):796-802- pdf


Friston KJ, Price CJ, Fletcher P, Moore C, Frackowiak RS, Dolan RJ.
The trouble with cognitive subtraction. 
NeuroImage. 1996 Oct;4(2):97-104.- pdf
Friston KJ 
Functional and effective connectivity in neuroimaging: A synthesis 
Human Brain Mapping 1995: 2;56-78- pdf
Friston KJ, Frith CD, Passingham RE, Liddle PF, Frackowiak RS.
Motor practice and neurophysiological adaptation in the cerebellum: a positron tomography study.
Proc Biol Sci. 1992 Jun 22;248(1323):223-8- pdf

Voxel-based morphometry and computational anatomy

Ashburner J, Friston KJ.
Unified segmentation. 
NeuroImage. 2005 Jul 1;26(3):839-51.- pdf
Friston KJ, Ashburner J.
Generative and recognition models for neuroanatomy. 
NeuroImage. 2004 Sep;23(1):21-4. 
Salmond CH, Ashburner J, Vargha-Khadem F, Connelly A, Gadian DG, Friston KJ.
Distributional assumptions in voxel-based morphometry. 
NeuroImage. 2002 Oct;17(2):1027-30.- pdf
Ashburner J, Friston KJ.
Voxel-based morphometry--the methods. 
NeuroImage. 2000 Jun;11(6 Pt 1):805-21.- pdf
Wright IC, McGuire PK, Poline JB, Travere JM, Murray RM, Frith CD, Frackowiak RS, Friston KJ.
A voxel-based method for the statistical analysis of gray and white matter density applied to schizophrenia. 
NeuroImage. 1995 Dec;2(4):244-52.- pdf
Friston KJ Ashburner J Frith CD Poline J-B Heather JD and Frackowiak RSJ 
The spatial registration and normalisation of images - pdf
Human Brain Mapping1995;3:165-189
Friston KJ, Passingham RE, Nutt JG, Heather JD, Sawle GV, Frackowiak RS.
Localisation in PET images: direct fitting of the intercommissural (AC-PC) line. 
J Cereb Blood Flow Metab. 1989 Oct;9(5):690-5. 

Hemodynamic models


J.M. Kilner, J. Mattout, R. Henson and K.J. Friston
Hemodynamic correlates of EEG: A heuristic
NeuroImage, Volume 28, Issue 1, 15 October 2005, Pages 280-286.- pdf
Mechelli A, Price CJ, Friston KJ.
Nonlinear coupling between evoked rCBF and BOLD signals: a simulation study of hemodynamic responses. 
NeuroImage. 2001 Oct;14(4):862-72.- pdf

Friston KJ, Mechelli A, Turner R, Price CJ

Nonlinear Responses in fMRI: The Balloon Model, Volterra Kernels and Other Hemodynamics

NeuroImage 12, 466-477 (2000)- pdf

Friston KJ, Josephs O, Rees G, Turner R.
Nonlinear event-related responses in fMRI. 
Magn Reson Med. 1998 Jan;39(1):41-52.- pdf
K.J. Friston, P. Jezzard, and R. Turner
Analysis of Functional MRI Time-Series 

Human Brain Mapping 1:153-171(1994)- pdf


Friston KJ

Regulation of rCBF by Diffusible Signals: An Analysis of Constraints on Diffusion and Elimination

Human Brain Mapping 3:56-65(1995)- pdf


Multivariate models

Friston KJ, Kahan J, Razi A, Stephan KE, Sporns O.
On nodes and modes in resting state fMRI.
Neuroimage. 2014 Oct;99:533-47.
Friston K, Chu C, Mourão-Miranda J, Hulme O, Rees G, Penny W, Ashburner J.
Bayesian decoding of brain images.
Neuroimage. 2007 Aug 24.- pdf
Friston K, Phillips J, Chawla D, Büchel C.
Nonlinear PCA: characterizing interactions between modes of brain activity. 
Philos Trans R Soc Lond B Biol Sci. 2000 Jan 29;355(1393):135-46.- pdf
Friston KJ, Frith CD, Fletcher P, Liddle PF, Frackowiak RS.
Functional topography: multidimensional scaling and functional connectivity in the brain.
Cereb Cortex. 1996 Mar-Apr;6(2):156-64.- pdf
Friston KJ, Frith CD, Frackowiak RS, Turner R.
Characterizing dynamic brain responses with fMRI: a multivariate approach.
Neuroimage. 1995 Jun;2(2):166-72.- pdf
Friston KJ, Frith CD, Liddle PF, Frackowiak RS.
Functional connectivity: the principal-component analysis of large (PET) data sets. 
J Cereb Blood Flow Metab. 1993 Jan;13(1):5-14.- pdf
K.J. Friston, L.G. Ungerleider, P. Jezzard, and R. Turner.
Characterizing modulatory interactions between V1 and V2 in human cortex with fMRI
Human Brain Mapping, 2:211-224, 1995. - pdf

Topological inference

Chumbley J, Worsley K, Flandin G, Friston K.
Topological FDR for neuroimaging.
Neuroimage. 2010 Feb 15;49(4):3057-64.- pdf
Friston KJ.
Testing for anatomically specified regional effects. 
Hum Brain Mapp. 1997;5(2):133-6.- pdf
Friston KJ, Holmes A, Poline JB, Price CJ, Frith CD.
Detecting activations in PET and fMRI: levels of inference and power. 
NeuroImage. 1996 Dec;4(3 Pt 1):223-35.- pdf
Worsley KJ Marrett S Neelin P Vandal AC Friston KJ Evans AC
A unified statistical approach for determining significant signals in images of cerebral activation
Human Brain Mapping1996;4:58-73.- pdf

Friston KJ Holmes AP Worsley KJ Poline JB Frith CD and Frackowiak RSJ

Statistical Parametric Maps in functional imaging: A general linear approach

Human Brain Mapping 1995;2;189-210.- pdf


Friston KJ Worsley KJ Frackowiak RSJ Mazziotta JC & Evans AC 
Assessing the significance of focal activations using their spatial extent 
Human Brain Mapping 1994;1:214-220.- pdf
Friston KJ, Frith CD, Liddle PF, Frackowiak RS.
Comparing functional (PET) images: the assessment of significant change. 
J Cereb Blood Flow Metab. 1991 Jul;11(4):690-9. - pdf
Friston KJ, Frith CD, Liddle PF, Dolan RJ, Lammertsma AA, Frackowiak RS. 
The relationship between global and local changes in PET scans. 
J Cereb Blood Flow Metab. 1990 Jul;10(4):458-66. - pdf



Source reconstruction (EEG and MEG)


Litvak V, Friston K.
Electromagnetic source reconstruction for group studies.
Neuroimage. 2008 Oct 1;42(4):1490-8.- pdf
Friston K, Harrison L, Daunizeau J, Kiebel S, Phillips C, Trujillo-Barreto N, Henson R, Flandin G, Mattout J.
Multiple sparse priors for the M/EEG inverse problem.
Neuroimage. 2007 Oct 10; [Epub ahead of print]- pdf
Mattout, J., Henson, R N. & Friston, K.J. 
Canonical Source Reconstruction for MEG.
Computational Intelligence and Neuroscience. 2007. Article ID 67613- pdf

Daunizeau J, Friston KJ.

A mesostate-space model for EEG and MEG.

NeuroImage. 2007 Jul 24;38(1):67-81- pdf


Friston K, Henson R, Phillips C, Mattout J.

Bayesian estimation of evoked and induced responses.

Hum Brain Mapp. 2006 Feb 1;27(9):722-735.- pdf


Mattout J, Phillips C, Penny WD, Rugg MD, Friston KJ.

MEG source localization under multiple constraints: An extended Bayesian framework.

NeuroImage. 2005 Dec 17; [Epub ahead of print]- pdf


Phillips C, Mattout J, Rugg MD, Maquet P, Friston KJ.
An empirical Bayesian solution to the source reconstruction problem in EEG. 
NeuroImage. 2005 Feb 15;24(4):997-1011.- pdf






Special Presentations




Special lecture: Free Energy and Active Inference - Karl J. Friston


Dynamic Causal Modelling- (PowerPoint)


Neural Mass Modelling- (PowerPoint)


Free-energy and the brain- (PowerPoint)


Free-energy and value- (PowerPoint)


Free-energy and precision- (PowerPoint)


Free-energy and saccades- (PowerPoint)


Free-energy and schizophrenia- (PowerPoint)


Cortical connectivity- (PowerPoint)


Variational filtering and DEM- (PowerPoint)


Free-energy lecture- (PowerPoint)


Predictive coding lecture- (PowerPoint)


Predictive coding and birdsong- (PowerPoint)


Brain connectivity- (PowerPoint)



Curriculum Vitae



Presentation I - (PowerPoint)


Presentation II - (PowerPoint)


Presentation III - (PowerPoint)